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One well-characterized non-histone substrate is HMGB1, a chromatin-binding protein with extracellular cytokine-like activity

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(10) I.T.Gajger et al (2018) Stable gastric pentadecapeptide BPC 157 in honeybee (Apis mellifera) therapy, to control Nosema ceranae invasions in apiary conditions Journal of Veterinary Pharmacology and Therapeutics, 2018 Aug;41(4):614-621